{"id":3399,"date":"2015-04-15T12:35:11","date_gmt":"2015-04-15T09:35:11","guid":{"rendered":"http:\/\/80.69.171.23\/py\/?page_id=3399"},"modified":"2026-01-14T10:17:22","modified_gmt":"2026-01-14T08:17:22","slug":"tutkimusjulkaisuja","status":"publish","type":"page","link":"https:\/\/paliskunnat.fi\/py\/organisaatio\/koeporotarha\/tutkimusjulkaisuja\/","title":{"rendered":"Tutkimusjulkaisuja"},"content":{"rendered":"

Kutuharjun tutkimusjulkaisuja<\/h2>\n

Vuosi 2025:<\/strong><\/p>\n

Vetter-Lang, S.G., Huber, N., Loe, L.E., Evans, A.L., Kumpula, J., Medb\u00f8e Thorsby, P. Ropstad, E. & Trondrud, L.M. Getting used to it? Stress of repeated management procedures in semidomesticated reindeer. BMC Veterinary Research (2025) 21: 268.
\nhttps:\/\/doi.org\/10.1186\/s12917-025-04718-8<\/a><\/p>\n

Vuosi 2024:<\/strong><\/p>\n

Boucher, A.J., Weladji, R.B., Holand, \u00d8., Kumpula, J. 2024: Modelling reindeer rut activity using on-animal acoustic recorders and machine learning. Ecology and Evolution. 2024;14:e11479.
\nhttps:\/\/doi.org\/10.1002\/ece3.11479<\/a><\/p>\n

Puch, L., Weladji, R. B., Holand, \u00d8., & Kumpula, J. 2024: The content of reindeer male vocalisations: acoustic cues to age and size. Bioacoustics, 33(4), 387\u2013399.
\nhttps:\/\/doi.org\/10.1080\/09524622.2024.2366924<\/a><\/p>\n

Vuosi 2023:<\/strong><\/p>\n

Trondrud, M., Pigeon, G., Kr\u00f3l, E., Albon, S. Ropstad, E., Kumpula, J., Evans, A.L., Speakman, J.R., Loe, L.E. 2023: A Summer Heat Wave Reduced Activity, Heart Rate, and Autumn Body Mass in a Cold-Adapted Ungulate. Physiological and Biochemical Zoology 96(4): s. 1-12.
\nhttps:\/\/doi.org\/10.1086\/725363<\/a><\/p>\n

Manni, K., Alenius, V., Kumpula, J., Pekkarinen, A.-J., Siitari, J. ja Tervonen, M. 2023: Kutuharjun koeporotarhan kehitt\u00e4minen \u2013 Tutkimustietoa tulevaisuudessakin elinkeinolle. Luonnonvara- ja biotalouden tutkimus 25\/2023. 48 sivua.
\nhttps:\/\/jukuri.luke.fi\/server\/api\/core\/bitstreams\/df72125d-61ae-4ef9-a639-31c5e14fa6f7\/content<\/a><\/p>\n

Vuosi 2022:<\/strong><\/p>\n

Manni, K., Kumpula, J., Pekkarinen, A., Alenius, V., Tervonen, M. ja Ollila, A. 2022. Kutuharjun koeporotarhan kehitt\u00e4missuunnitelma valmistumassa. Poromies 91(4): 55-57.<\/p>\n

Vuosi 2021:<\/strong><\/p>\n

Nagy, S., Lindeberg, H., Nikitkina, E., Krutikova, A., Smith, E., Kumpula, J. & Holand, \u00d8. 2021: Reproduction of male reindeer (Rangifer tarandus). Animal Reproduction Science 227 (2021) 106722, 1-8 p.
\nhttps:\/\/doi.org\/10.1016\/j.anireprosci.2021.106722<\/a><\/p>\n

Lindeberg H., Nikitkina, E., Nagy, Sz., Musidray, A., Shiryaev, G., Kumpula, J. & Holand, \u00d8 2021: Potential applications of assisted reproductive technologies (ART) in reindeer (Rangifer tarandus). Animal Reproduction Science 235 (2021) 106890.
\nhttps:\/\/doi.org\/10.1016\/j.anireprosci.2021.106890<\/a><\/p>\n

Driscoll, J.G., Alo, F.M., Paoli, A., Weladji, R.B., Holand, \u00d8., Kumpula, J. & Soveri T. 2021. Influence of operational sex ratio and male age on mating competition intensity in reindeer (Rangifer tarandus). Ethology Ecology & Evolution, 2021.
\nhttps:\/\/doi.org\/10.1080\/03949370.2021.1989054<\/a><\/p>\n

Alenius, V., Kumpula, J., Manni, K., Pekkarinen, A., Ollila A. ja Tervonen, M. 2021. Selvitysty\u00f6 Kutuharjun koeporotarhan tutkimusinfran uudistamiseksi alkanut. Poromies 90(2): 50-51.<\/p>\n

Vuosi 2020:<\/strong><\/p>\n

Holand, H., Kvalnes, T., R\u00f8ed, K.H., Holand, \u00d8., S\u00e6ther B.-E. & Kumpula, J. 2020. Stabilizing selection and adaptive evolution in a combination of two traits in an arctic ungulate. Evolution 74(1): 103\u2013115. https:\/\/doi.org\/10.1111\/evo.13894<\/a><\/p>\n

Paoli, A., Weladji, R.B., Holand, \u00d8. & Kumpula, J. 2020. The onset in spring and the end in autumn of the thermal and vegetative growing season affect calving time and reproductive succhttps:\/\/doi:10.1111\/evo.13894ess in reindeer. Current Zoology, Current Zoology, 2020, 66(2), 123\u2013134 (online version first published in Current Zoology 2019, 1\u201312.
\nhttps:\/\/doi.org\/10.1093\/cz\/zoz032<\/a>)<\/p>\n

Paoli, A., Weladji, R.B., Holand, \u00d8. & Kumpula, J. 2020. Response of reindeer mating time to climatic variability. BMC Ecology 20(44): 1-13.
\nhttps:\/\/doi.org\/10.1186\/s12898-020-00312-8<\/a>.<\/p>\n

de Villemereuil, P., Charmantier, A., Arlt, A., Bize, P., Brekke, P., Brouwer, L., Cockburn, A., C\u00f4t\u00e9, S. D., Dobson, F. S., Evans, S. R., Festa-Bianchet, M., Gamelon, M., Hamel, S., Hegelbach, J., Jerstad, K., Kempenaers, B., Kruuk, L. E. B., Kumpula, J., Kvalnes, T., McAdam,, A. G., McFarlane, S. E., Morrissey, M. B., P\u00e4rt, T., Pemberton, J. M., Qvarnstr\u00f6m, A., R\u00f8stad, O-W., Schroeder, J., Senar, J. C., Sheldon, B. C., van de Pol, M., Visser, M. E., Wheelwright, N. T., Tufto, J., & Chevin, L-M., 2020. Fluctuating optimum and temporally variable selection in the wild. PNAS 117 (50) 31969-31978; first published online:
\nhttps:\/\/doi.org\/10.1073\/pnas.2009003117<\/a><\/p>\n

Vuosi 2019:<\/strong><\/p>\n

Paoli, A., Weladji, R.B., Holand, \u00d8. & Kumpula, J. 2019. Early\u2010life conditions determine the between\u2010individual heterogeneity in plasticity of calving date in reindeer. Journal of Animal Ecology 2019: 1\u201314.
\nhttps:\/\/doi.org\/10.1111\/1365-2656.13096<\/a><\/p>\n

Paoli, A. 2019. Breeding phenology of a semi-domesticated reindeer (Rangifer tarandus) population in response to climatic variability. PhD thesis, Concordia University, Canada.<\/p>\n

Vuosi 2018:<\/strong><\/p>\n

Holand, \u00d8. & Weladji, R. B. 2018. Rangifer Mating Strategies. In Tryvan, M, Kutz, SJ (eds.) Reindeer and Caribou: Health and Disease. CRC Press, Boca Raton, London, New York.<\/p>\n

Paoli, A, Weladji, R. B., Holand, \u00d8. & Kumpula, J. 2018. Winter and spring climatic conditions influence timing and synchrony of calving in reindeer. PLoS ONE 13(4): e0195603.
\nhttps:\/\/doi.org\/10.1371\/journal.pone.0195603<\/a><\/p>\n

Vuosi 2017:<\/strong><\/p>\n

Strong, J. S., Weladji, R. B., Holand, \u00d8., R\u00f8ed, K. H. & Nieminen, M. 2017. Personality and fitness consequences of flight initiation distance and mating behavior in subdominant male reindeer (Rangifer tarandus). Ethology 123: 484\u2013492.<\/p>\n

Weladji, R. B., Body, G., Holand, \u00d8., Meng, X. & Nieminen, M. 2017. Temporal variation in the operational sex ratio and male mating behaviours in reindeer (Rangifer tarandus). Behavioural Processes 140:96-103.<\/p>\n

Vuosi 2016:<\/strong><\/p>\n

Engelhardt, S. C., Weladji, R. B., Holand, \u00d8., R\u00f8ed, K. H. & Nieminen, M. 2016. Allonursing in reindeer, Rangifer tarandus: a test of the kin-selection hypothesis. Journal of Mammalogy 97: 689-700. DOI: http:\/\/dx.doi.org\/10.1093\/jmammal\/gyw027<\/p>\n

Engelhardt, S. C., Weladji,\u00a0 R. B., Holand, \u00d8. & Nieminen, M. 2016. Allosuckling in reindeer (Rangifer tarandus): A test of the improved nutrition and compensation hypotheses. Mammalian Biology 81:146-152<\/p>\n

Vuosi 2015:<\/strong><\/p>\n

Body, G., Weladji, R. B., Holand, \u00d8. & Nieminen, M. 2015. Measuring variation in the frequency of group fission and fusion from continuous monitoring of group sizes. J. Mammal. 94: 791-799.<\/p>\n

Body, G., Weladji, R. B., Holand,\u00a0 \u00d8. & Nieminen, M. 2015. Fission-fusion group dynamics in reindeer reveal an increase of cohesiveness at the beginning of the peak rut. Acta Ethologica 18:101-110<\/p>\n

Djakovic, N., Holand, \u00d8., Hovland, A. L., R\u00f8ed, K. H., Weladji, R. B. & Nieminen, M. 2015. Effects of males’ presence on female behaviour during the rut. Ethology Ecology & Evolution 27:148-160.<\/p>\n

Engelhardt, S. C., Weladji,\u00a0 R. B., Holand, \u00d8., R\u00f8ed, K. H. & Nieminen, M. 2015. Evidence of Reciprocal Allonursing in Reindeer, Rangifer tarandus. Ethology 121:245-259<\/p>\n

Pintus, E., Uccdeddu, S., R\u00f8ed, K. H., Gonzalez,\u00a0 J. P., Carranza, J., Nieminen,\u00a0 M. & Holand, \u00d8. 2015. Flexible mating tactics and associated reproductive effort during the rutting season in male reindeer (Rangifer tarandus, L. 1758). Current Zool. 61:802-810.<\/p>\n

Uccheddu, S., Body, G., Weladji, R. B., Holand, \u00d8. & Nieminen, M. 2015. Foraging competition in larger groups overrides harassment avoidance benefits in female reindeer (Rangifer tarandus). Oecologia\u00a0 179: 711-718. DOI 10.1007\/s00442-015-3392-5<\/p>\n

Vuosi 2014:<\/strong><\/p>\n

Body, G., Weladji, R. B., Holand, \u00d8. & Nieminen, M. 2014. Fission-fusion group dynamics in reindeer reveal an increase of cohesiveness at the beginning of the peak rut. Acta ethol. DOI: 10.1007\/S10211-014-0190-8<\/p>\n

Body, G., Weladji, R. B., Holand, \u00d8. & Nieminen, M. 2014. Highly Competitive Reindeer Males Control Female Behavior during the Rut. PLoS one 9. DOI: 10.1371\/journal.pone.0095618<\/p>\n

Engelhardt, S., Holand, \u00d8., Nieminen, M. & Weladji, R. B. 2014. Allosuckling in reindeer (Rangifer tarandus): milk-theft, mismothering or kin selection?”\u00a0 Behavioural Processes: 107:133-141<\/p>\n

Holand, \u00d8. 2014. Reinens reproduksjonsstrategi (in Norwegian). Norsk Vet Tidskrift, 2:92-100.<\/p>\n

Muuttoranta, K. & M\u00e4ki-Tanila, A. 2014. Reindeer production in Finland \u2013 integration of traditional and new technologies. Kappale kirjassa: Cattle husbandry in Eastern Europe and China – Structure, development paths and optimization. Toim.: Kuipers, A., Roztalnyy, A. and Keane, G. EAAP Scientific Series, Publication No. 135. ISBN: 978-90-8686-232-0978, DOI: 10.3920\/978-90-8686-785-1.<\/p>\n

V\u00e5ge, D. I., Nieminen, M., Andersen, D. & R\u00f8ed, K. H. 2014 Two missense mutations in melanocortin 1 receptor (MC1R) are strongly associated with dark ventral coat color in reindeer (Rangifer tarandus). Animal Genetics 45 (5): 750-753<\/p>\n

Vuosi 2013:<\/strong><\/p>\n

Melnycky, N. A., Weladji,\u00a0 R. B., Holand,\u00a0 \u00d8. & Nieminen, M. 2013. Scaling of antler size in reindeer (Rangifer tarandus): sexual dimorphism and variability in resource allocation. Journal of Mammalogy 94: 1371-1379.<\/p>\n

Muuttoranta, K., Tapio, M., Holand, \u00d8., R\u00f8ed, K. H., Nieminen, M. & M\u00e4ki-Tanila, A. 2013. Genetic and environmental effects in the variation of birth date and birth weight of reindeer calves. Rangifer 33(1):25-35. http:\/\/dx.doi.org\/10.7557\/2.33.1.2598<\/p>\n

Muuttoranta, K., Tapio, M., Holand, \u00d8., R\u00f8ed, K. H., Nieminen, M. & M\u00e4ki-Tanila, A. 2013. Genetic variation in meat production related traits in reindeer (Rangifer t. tarandus). Rangifer 34: 21-36.<\/p>\n

Vuosi 2012:<\/strong><\/p>\n

Muuttoranta, K. & M\u00e4ki-Tanila, A. 2012. Regional differences in reindeer herding operations in Finland. Acta Agricultur\u00e6 Scandinavica, Section A \u2013 Animal Science,62(3): 142 \u2013 152. DOI:10.1080\/09064702.2012.747560<\/p>\n

Muuttoranta, K. & M\u00e4ki-Tanila, A. 2012. The reindeer herders are interested in selective breeding. Presentation in the 63rd Annual Meeting of the European Federation of Animal Science, Bratislava, Slovakia, 27 \u2013 31 August 2012. Book of abstracts No 18 (2012).<\/p>\n

Holand, \u00d8., L\u2019Italien, L., Weladji, R. B., Djakovi\u0107, N., R\u00f8ed, K., Hovland, A. L. & Nieminen, M. 2012. Shit happens \u2013 a glimpse into males\u2019 mating tactics in a polygynous ungulate – the reindeer. Rangifer 32: 65-72.<\/p>\n

Body, G., Weladji, R. B. & Holand, \u00d8. 2012. The recursive model as a new approach to validate and monitor activity sensors. Behavioral Ecology and Sociobiology 66: 1531-1541<\/p>\n

L\u2019Italien, L., Weladji, R. B., Holand, \u00d8., R\u00f8ed, K. H., Nieminen, M. & C\u00f4t\u00e9, S. D. 2012. Mating group size and stability in reindeer: the effects of male characteristics, sex ratio and male age-structure. Ethology 118: 783-792<\/p>\n

Holand, \u00d8., Weladji, R. B., Mysterud, A., R\u00f8ed, K. H., Reimers, E. & Nieminen, M. 2012. Induced orphaning reveals post-weaning maternal care in reindeer. European Journal of Wildlife Research 58: 589-596.<\/p>\n

Tennenhouse, E. M., Weladji, R. B., Holand, \u00d8. & Nieminen, M. 2012. Timing of reproductive effort differs between young and mature male reindeer. Annales Zoologici Fennici 49: 152-160.<\/p>\n

Djakovi\u0107, N., Holand, \u00d8., Hovland, A. L., R\u00f8ed, K., Weladji, R. B., Fjelstad, E. & Nieminen, M. 2012. Association patterns and kinship in female reindeer (Rangifer tarandus) during rut. Acta Ethologica\u00a0 15: 165\u2013171<\/p>\n

Vuosi 2011:<\/strong><\/p>\n

Muuttoranta, K. & M\u00e4ki-Tanila, A. 2011. Selection decisions among reindeer herders in Finland. Rangifer 31(1): 129 \u2013 138.<\/p>\n

Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2011. Influence of dam information on selection practices in reindeer.\u00a0 Presentation in the 13th International Arctic Ungulates Conference, Yellowknife, Northwest Territories, Canada, 22\u201326 August 2011.<\/p>\n

Morden, C. C., Weladji, R. B., Ropstad, E., Dahl E., Holand, \u00d8., Mastromonaco, G. & Nieminen, M. 2011. Faecal hormones as a non-invasive population monitoring method for reindeer and caribou. Journal of Wildlife Management 75: 1426-1435.<\/p>\n

Tennenhouse, E. M., Weladji, R. B., Holand, \u00d8., R\u00f8ed, K. H. & Nieminen, M. 2011. Mating group composition influence somatic costs and activity in rutting dominant male reindeer (Rangifer tarandus). Behavioral Ecology and Sociobiology 65: 287-295<\/p>\n

Vuosi 2010 (t\u00e4st\u00e4 eteenp\u00e4in kaikki julkaisut mukana):<\/strong><\/p>\n

Muuttoranta, K. & M\u00e4ki-Tanila, A. 2010. Haastattelututkimus porojen valinnan tavoitteista. Poromies 5\/2010: 44\u201345.<\/p>\n

Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2010. Em\u00e4ll\u00e4 on suuri vaikutus vasan kasvuun. Poromies 5\/2010: 45\u201346.<\/p>\n

Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2010. Estimating Maternal Effects on Growth of Reindeer (Rangifer t. tarandus). Proceedings of the 9th World Congress on Genetics Applied to Livestock Production, Leipzig, Germany, 1st \u2013 6th August 2010. ISBN 978-3-00-031608-1<\/p>\n

M\u00e4ki-Tanila, A., Muuttoranta, K. & Nieminen, M. 2010. Maternal effects affect the genetic variation of meat production in reindeer. Presentation in the 61st Annual Meeting of the European Association for Animal Production, Heraklion, Greece, 23 \u2013 27 August 2010<\/p>\n

Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2010. Estimating maternal effects on weight records of Kutuharju reindeer. Presentation in the 16th Nordic Conference on Reindeer and Reindeer Husbandry Research, Troms\u00f8, Norway, 16 \u2013 18 November 2010. Rangifer Report No. 14 (2010), p. 103-104.<\/p>\n

Weladji, R. B., Holand, \u00d8., Gaillard, J-M., Yoccoz, N. G., Mysterud, A., Nieminen, M. & Stenseth, N. 2010. Age-specific changes in different components of reproductive output in female reindeer: Terminal allocation or senescence? Oecologia 162: 261-271<\/p>\n

Vuosi 2009 (ei norjalaisia tai kanadalaisia):<\/strong><\/p>\n

Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2009. Em\u00e4t vaikuttavat vasoihin pitk\u00e4\u00e4n. Poromies 5\/2009: 30.<\/p>\n

Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2009. El\u00e4invalinnalla enemm\u00e4n euroja. Poromies 6\/2009: 24\u201325.<\/p>\n

Porop\u00e4ivien esityksi\u00e4 (vain Muuttoranta ym. pit\u00e4m\u00e4t listattu) <\/strong><\/p>\n

24-25.4.2008: Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2008. Kutuharjun porot soveltuvat hyvin genetiikkatutkimukseen.<\/p>\n

23-24.4. 2009: Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2009. Em\u00e4t vaikuttavat vasoihin pitk\u00e4\u00e4n.<\/p>\n

22- 23.4.2010: Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2010. Em\u00e4ll\u00e4 on suuri vaikutus vasan kasvuun.<\/p>\n

25-26.4.2013: Muuttoranta, K. & M\u00e4ki-Tanila, A. 2013. Valinta on t\u00e4rke\u00e4 porotalouden elementti.<\/p>\n

25-26.4.2013: Muuttoranta, K., Nieminen, M. & M\u00e4ki-Tanila, A. 2013. Vasaominaisuuksien vaihtelun p\u00e4\u00e4tekij\u00e4t.<\/p>\n

 <\/p>\n

 <\/p>\n","protected":false},"excerpt":{"rendered":"

Kutuharjun tutkimusjulkaisuja Vuosi 2025: Vetter-Lang, S.G., Huber, N., Loe, L.E., Evans, A.L., Kumpula, J., Medb\u00f8e Thorsby, P. Ropstad, E. & Trondrud, L.M. Getting used to it? Stress of repeated management procedures in semidomesticated reindeer. BMC Veterinary Research (2025) 21: 268. https:\/\/doi.org\/10.1186\/s12917-025-04718-8 Vuosi 2024: Boucher, A.J., Weladji, R.B., Holand, \u00d8., Kumpula, J. 2024: Modelling reindeer rut […]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":311,"menu_order":2,"comment_status":"open","ping_status":"closed","template":"","meta":{"footnotes":""},"class_list":["post-3399","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/pages\/3399","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/comments?post=3399"}],"version-history":[{"count":39,"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/pages\/3399\/revisions"}],"predecessor-version":[{"id":8727,"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/pages\/3399\/revisions\/8727"}],"up":[{"embeddable":true,"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/pages\/311"}],"wp:attachment":[{"href":"https:\/\/paliskunnat.fi\/py\/wp-json\/wp\/v2\/media?parent=3399"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}